TITLE t-calcium channel
: t-type calcium channel
UNITS {
(mA) = (milliamp)
(mV) = (millivolt)
FARADAY = 96520 (coul)
R = 8.3134 (joule/degK)
KTOMV = .0853 (mV/degC)
}
INDEPENDENT {t FROM 0 TO 1 WITH 1 (ms)}
PARAMETER {
dt (ms)
v (mV)
tBase = 23.5 (degC)
celsius = 22 (degC)
gcatbar = 0 (mho/cm2)
ki=.001 (mM)
cai=5.e-5 (mM)
cao = 2 (mM)
eca = 140
VhalfH= -85
VhalfM= -47
gh = 0.4
zh = 5
gm = 0.40
zm = 5
aoh = 0.3
aom = 0.03
tfi = 10
tfa = 1
k
tadj
taum
tauh
}
NEURON {
SUFFIX mycat
USEION ca READ cai,cao WRITE ica
: USEION Ca WRITE ica VALENCE 2
: The T-current does not activate calcium-dependent currents.
: The construction with dummy ion Ca prevents the updating of the
: internal calcium concentration.
RANGE gcatbar, hinf, minf, taum, tauh, ica
: RANGE gcatbar, hinf, minf, iCa
}
STATE {
m h
}
ASSIGNED {
ica (mA/cm2)
gcat (mho/cm2)
minf
hinf
}
INITIAL {
k = 1000*8.3134*(celsius+273.15)/96520
tadj = 3^((celsius-tBase)/10) : assume Q10 of 3
rates(v)
m = minf
h = hinf
gcat = gcatbar*m*m*h*h2(cai)
}
BREAKPOINT {
SOLVE states
gcat = gcatbar*m*m*h*h2(cai)
ica = gcat*ghk(v,cai,cao)
}
UNITSOFF
FUNCTION h2(cai(mM)) {
h2 = ki/(ki+cai)
}
FUNCTION ghk(v(mV), ci(mM), co(mM)) (mV) {
LOCAL nu,f
f = KTF(celsius)/2
nu = v/f
ghk=-f*(1. - (ci/co)*exp(nu))*efun(nu)
}
FUNCTION KTF(celsius (degC)) (mV) {
KTF = ((25./293.15)*(celsius + 273.15))
}
FUNCTION efun(z) {
if (fabs(z) < 1e-4) {
efun = 1 - z/2
}else{
efun = z/(exp(z) - 1)
}
}
FUNCTION alph(v(mV)) {
TABLE FROM -150 TO 150 WITH 200
alph = aoh*exp(-gh*zh*(v-VhalfH)/k)
}
FUNCTION beth(v(mV)) {
TABLE FROM -150 TO 150 WITH 200
beth = (aoh/10)*exp((1-gh)*zh*(v-VhalfH)/k)
}
FUNCTION alpm(v(mV)) {
TABLE FROM -150 TO 150 WITH 200
alpm = aom*exp((1-gm)*zm*(v-VhalfM)/k)
}
FUNCTION betm(v(mV)) {
TABLE FROM -150 TO 150 WITH 200
betm = aom*exp(-gm*zm*(v-VhalfM)/k)
}
UNITSON
LOCAL facm,fach
:if state_cagk is called from hoc, garbage or segmentation violation will
:result because range variables won't have correct pointer. This is because
: only BREAKPOINT sets up the correct pointers to range variables.
PROCEDURE states() { : exact when v held constant; integrates over dt step
rates(v)
m = m + facm*(minf - m)
h = h + fach*(hinf - h)
VERBATIM
return 0;
ENDVERBATIM
}
PROCEDURE rates(v (mV)) { :callable from hoc
LOCAL a
a = alpm(v)
: taum = 10/(a + betm(v))
taum = 20
minf = a/(a+betm(v))
facm = (1 - exp(-dt/taum))
a = alph(v)
: tauh = 1000/(a + beth(v))
tauh = 60
hinf = a/(a+beth(v))
fach = (1 - exp(-dt/tauh))
}