TITLE calcium channels (L, N, and T types)
UNITS {
(molar) = (1/liter)
(S) = (siemens)
(mA) = (milliamp)
(mV) = (millivolt)
(mM) = (millimolar)
F = (faraday) (coulomb)
R = (mole k) (mV-coulomb/degC)
}
NEURON {
SUFFIX cachan
USEION ca READ cai WRITE ica
RANGE gcalbar,gcanbar,gcatbar,ica,ical,icat,ican,kml,kmn
GLOBAL dlinf,dninf,dtinf,ftinf,flinf,fninf
}
INDEPENDENT {t FROM 0 TO 1 WITH 1 (ms)}
PARAMETER {
v (mV)
dt (ms)
cai (mM)
celsius = 35.0 (degC)
gcatbar = 1044.0e-6 (S/cm2)
gcanbar = 171.0e-6 (S/cm2)
gcalbar = 216.0e-6 (S/cm2)
kmn = 0.001 (mM)
kml = 0.00045 (mM)
eca = 120 (mV)
cao = 2.0 (mM)
}
STATE {
d_t dl dn ft
}
ASSIGNED {
ica (mA/cm2)
ical (mA/cm2)
ican (mA/cm2)
icat (mA/cm2)
dlinf dninf dtinf ftinf fninf flinf
}
BREAKPOINT {
SOLVE states METHOD cnexp
fninf = kmn/(kmn + cai)
flinf = kml/(kml + cai)
ical = gcalbar*dl*flinf*(v - eca)
ican = gcanbar*dn*fninf*(v - eca)
icat = gcatbar*d_t*ft*(v - eca)
ica = ical + ican + icat
}
UNITSOFF
INITIAL {
dl = boltz(v,-50.0,3.0)
dn = boltz(v,-45.0,7.0)
d_t = boltz(v,-63.5,1.5)
ft = boltz(v,-76.2,-3.0)
}
DERIVATIVE states { :Computes state variables m, h, and n
LOCAL dlinf,dninf,dtinf,ftinf,dltau,dntau,dttau,fttau
dlinf = boltz(v,-50.0,3.0)
dninf = boltz(v,-45.0,7.0)
dtinf = boltz(v,-63.5,1.5)
ftinf = boltz(v,-76.2,-3.0)
dltau = gaussian(v,18.0,20.0,45.0,1.50)
dntau = gaussian(v,18.0,25.0,70.0,0.30)
dttau = gaussian(v,65.0,6.32455,66.0,3.5)
fttau = gaussian(v,50.0,10.0,72.0,10.0)
dl' = (dlinf-dl)/dltau
dn' = (dninf-dn)/dntau
d_t' = (dtinf-d_t)/dttau
ft' = (ftinf-ft)/fttau
}
FUNCTION gaussian(v,a,b,c,d) {
LOCAL arg
arg= a*exp(-(c+v)*(v+c)/(b*b)) +d
gaussian = arg
}
FUNCTION boltz(x,y,z) {
LOCAL arg
arg= -(x-y)/z
if (arg > 50) {boltz = 0}
else {if (arg < -50) {boltz = 1}
else {boltz = 1.0/(1.0 + exp(arg))}}
}
UNITSON