//genesis // this is copied from the granule cell model by RM on 19 June 2007 /********************************************************************* ** The synaptic conductance equations *********************************************************************/ float Q10_synapse = 3.0 function make_Granule_syns //[Mg] in mM float CMg = 1.2 // per mM float eta = 0.2801 // per V float gamma = 62 float offset = - 0.01 echo eta = {eta} eta = eta * {exp {- gamma * offset}} echo new eta = {eta} echo diag Gran_syn1 0 /* NMDA channel made by CP */ /* From Gabbiani et al. (model) 1994 */ echo diag Gran_syn1 2 if (!({exists NMDA})) create synchan2 NMDA end setfield NMDA Ek {E_NMDA} tau2 {3e-3 / Q10_synapse} \ tau1 {40e-3 / Q10_synapse} \ gmax {G_NMDA} // use the following value for synaptic activation when TEST.g is run // gmax {4.0 * G_NMDA} if (! {exists NMDA/Mg_BLOCK}) create Mg_block NMDA/Mg_BLOCK end setfield NMDA/Mg_BLOCK CMg {CMg} \ KMg_A {1/eta} \ \\ *({exp {EREST_ACT*gamma}})} \ KMg_B {1.0/gamma} echo diag Gran_syn1 3 /* AMPA channel, made by CP */ /* Reference: voltage clamp data from ** Gabbiani et al. (model): J.Neurophys. 1994 ** Silver et al.: Nature 1992. */ if (!({exists AMPA})) create synchan2 AMPA end //sec //sec setfield AMPA Ek {E_AMPA} tau2 {0.09e-3 / Q10_synapse} \ tau1 {1.5e-3 / Q10_synapse} \ gmax {G_AMPA} // use the following value for synaptic activation when TEST.g is run // gmax {6.0 * G_AMPA} // Incorporate constant of proportionality (1.273) in G_AMPA echo diag Gran_syn1 4 /* GABAA? channel, made by EDS */ /* Reference: voltage clamp data from ** Miles R: . J Physiol 1990. ** Tpeak: 3.25 ms, Tdecay = 28 ms ** Note the high frequency. This is being used to model tonic ** Golgi cell inhibition to the granule cell. */ if (!({exists GABAA})) create synchan2 GABAA end //sec //sec // should be large for tonic inhibition setfield GABAA Ek {E_GABAA} tau1 {0.93e-3 / Q10_synapse} \ tau2 {26.50e-3 / Q10_synapse} \ gmax {G_GABAA} frequency {0.0} // setfield GABAA Ek {E_GABAA} tau1 {0.93e-3} \ // tau2 {26.50e-3} \ // gmax {G_GABAA} frequency {0.0} // use the following value for synaptic activation when TEST.g is run // gmax {G_GABAA * 45.0 * 0.6 / 14.14} frequency {0.0} /* GABA_B channel, using a dual exponential function with time constants of 80 ** and 40 msec as in Suarez, Koch and Douglas 1995 (J. Neurosci. 15, ** 6700-1719; cat visual cortex). ** A more detailed model can be found in Otis, De Koninck and Mody 1993 ** (J. Physiol. 463, 391-407; rat hippocampal slices; this model uses 4th ** power exponential activation and dual exponential inactivation). ** See also Benardo 1994 (J. Physiol. 476.2, 203-215; slice rat neocortex) ** and Connors, Malenka and Silva 1988 (J. Physiol. 406, 443-468; slice ** rat and cat visual cortex. */ if (!({exists GABAB})) create synchan2 GABAB end setfield GABAB Ek {E_GABAB} tau1 {0.080 / Q10_synapse} \ tau2 {0.040 / Q10_synapse} \ gmax {G_GABAB} frequency {0.0} end