TITLE I-h channel from Magee 1998 for distal dendrites
UNITS {
(mA) = (milliamp)
(mV) = (millivolt)
}
PARAMETER {
curr
v (mV)
ehd = -30 (mV)
celsius (degC)
ghdbar=.0001 (mho/cm2)
vhalfl=-81 (mV)
kl=-8
vhalft=-75 (mV)
a0t=0.011 (/ms)
zetat=2.2 (1)
gmt=.4 (1)
q10=4.5
qtl=1
sh=0 (mV)
time1=600
time0=100
alphash0=0
alphash1=0.13
alphahd=0.3
ehd2
sh2
count=1
vrun (mV)
delta=0
vinit=-76.2
vvrun=0
timestep=1000
vrun2
v0
dv0
ddv
flag=0
FNa = 2
PNa = 1
BNa = 2.6
CNa = 60
stim_moltNa=1
}
NEURON {
SUFFIX hd
POINTER stim_i
NONSPECIFIC_CURRENT i
RANGE flag, curr, ghdbar, vhalfl, sh,count,delta2,vrun2, stim_moltNa, ehd2
GLOBAL linf,taul
}
STATE {
l
}
ASSIGNED {
i (mA/cm2)
linf
taul
ghd
stim_i
}
INITIAL {
rate(v,sh2)
l=linf
vrun=0
vvrun=vrun
}
BREAKPOINT {
SOLVE states METHOD cnexp
ghd = ghdbar*l
ehd2=ehd-alphahd*vvrun
i = ghd*(v-ehd2)
}
FUNCTION alpt(v(mV)) {
alpt = exp(0.0378*zetat*(v-vhalft))
}
FUNCTION bett(v(mV)) {
bett = exp(0.0378*zetat*gmt*(v-vhalft))
}
DERIVATIVE states { : exact when v held constant; integrates over dt step
rate(v,sh2)
l' = (linf - l)/taul
}
BEFORE STEP { LOCAL i
if(stim_i==0 && flag==0){
vrun=0
vvrun=0
}else{
flag=1
delta=v-vinit
if (count<timestep+1){
vrun= (delta-vrun)*(FNa/(count+1))+vrun
vrun2=vrun
}else{
vrun2= (delta)*(FNa/(timestep+1))+vrun2*pow((1-FNa/(timestep+1)),PNa)
}
vvrun=(BNa*vrun2/(1+vrun2/CNa))
count=count+1
}
sh2=sh+alphash1*vvrun
}
PROCEDURE rate(v (mV),sh2) { :callable from hoc
LOCAL a,qt,i
qt=q10^((celsius-33)/10)
a = alpt(v-sh2)
linf = 1/(1 + exp(-(v-vhalfl-sh2)/kl))
: linf = 1/(1+ alpl(v-sh2))
taul = bett(v-sh2)/(qtl*qt*a0t*(1+a))
}