TITLE nax
: Na current for axon. No slow inact.
: M.Migliore Jul. 1997
: added sh to account for higher threshold M.Migliore, Apr.2002
NEURON {
SUFFIX nax
POINTER stim_i
USEION na READ ena WRITE ina
RANGE flag, curr, gbar, sh,ena2,alpha,vrun,count,vvrun,delta2,vrun2, stim_moltNa,flag
GLOBAL minf, hinf, mtau, htau,thinf, qinf,sh2
}
PARAMETER {
curr
sh = 0 (mV)
gbar = 0.010 (mho/cm2)
tha = -30 (mV) : v 1/2 for act
qa = 7.5 (mV) : act slope (4.5)
Ra = 0.4 (/ms) : open (v)
Rb = 0.124 (/ms) : close (v)
thi1 = -45 (mV) : v 1/2 for inact
thi2 = -45 (mV) : v 1/2 for inact
qd = 1.5 (mV) : inact tau slope
qg = 1.5 (mV)
mmin=0.02
hmin=0.5
q10=2
Rg = 0.01 (/ms) : inact recov (v)
Rd = .03 (/ms) : inact (v)
thinf = -53 (mV) : inact inf slope
qinf = 2 (mV) : inact inf slope
ena (mV) : must be explicitly def. in hoc
ena2 (mV)
celsius
v (mV)
vinit=-76.2 (mV)
delta=0
count=1
time0=100
time1=600
sh2
alphash0=0.1
alphash1=0.13
alphaena=0.25
count2=1
vvrun=0
timestep=1000
vrun2
v0
dv0
ddv
flag=0
FNa = 2
PNa = 1
BNa = 2.6
CNa = 60
stim_moltNa=1
}
UNITS {
(mA) = (milliamp)
(mV) = (millivolt)
(pS) = (picosiemens)
(um) = (micron)
}
ASSIGNED {
ina (mA/cm2)
thegna (mho/cm2)
minf hinf
mtau (ms) htau (ms)
vrun
stim_i
}
STATE { m h}
BREAKPOINT {
SOLVE states METHOD cnexp
thegna = gbar*m*m*m*h
ena2=ena-alphaena*vvrun
ina = thegna * (v - ena2)
}
INITIAL {
trates(v,sh2)
m=minf
h=hinf
vrun=0
vvrun=vrun
}
DERIVATIVE states {
trates(v,sh2)
m' = (minf-m)/mtau
h' = (hinf-h)/htau
}
BEFORE STEP { LOCAL i
if(stim_i==0 && flag==0){
vrun=0
vvrun=0
}else{
flag=1
delta=v-vinit
if (count<timestep+1){
vrun= (delta-vrun)*(FNa/(count+1))+vrun
vrun2=vrun
}else{
vrun2= (delta)*(FNa/(timestep+1))+vrun2*pow((1-FNa/(timestep+1)),PNa)
}
vvrun=(BNa*vrun2/(1+vrun2/CNa))
count=count+1
}
sh2=sh+alphash1*vvrun
}
PROCEDURE trates(vm,sh2) {
LOCAL a, b, qt,i
qt=q10^((celsius-24)/10)
a = trap0(vm,tha+sh2,Ra,qa)
b = trap0(-vm,-tha-sh2,Rb,qa)
mtau = 1/(a+b)/qt
if (mtau<mmin) {mtau=mmin}
minf = a/(a+b)
a = trap0(vm,thi1+sh2,Rd,qd)
b = trap0(-vm,-thi2-sh2,Rg,qg)
htau = 1/(a+b)/qt
if (htau<hmin) {htau=hmin}
hinf = 1/(1+exp((vm-thinf-sh2)/qinf))
}
FUNCTION trap0(v,th,a,q) {
if (fabs(v-th) > 1e-6) {
trap0 = a * (v - th) / (1 - exp(-(v - th)/q))
} else {
trap0 = a * q
}
}