COMMENT
//****************************//
// Created by Alon Polsky //
// apmega@yahoo.com //
// 2010 //
//****************************//
Modified 2015 by Robert Egger
to include facilitation variable
as modeled by Varela et al. 1997
ENDCOMMENT
TITLE NMDA synapse
NEURON {
POINT_PROCESS A1_IB
NONSPECIFIC_CURRENT inmda,iampa
RANGE gampamax,gnmdamax,inmda,iampa
RANGE gnmda,gampa
RANGE e,tau1,tau2,tau3,tau4
}
UNITS {
(nA) = (nanoamp)
(mV) = (millivolt)
(nS) = (nanomho)
(mM) = (milli/liter)
(mA) = (milliamp)
(um) = (micron)
}
PARAMETER {
gnmdamax=1 (nS)
gampamax=1 (nS)
e= 0.0 (mV)
tau1= 30 (ms): NMDA inactivation
tau2=4 (ms) : NMDA activation
tau3= 2 (ms) : AMPA inactivation
tau4=0.2 (ms) : AMPA activation
n=0.05 (/mM)
gama=0.05 (/mV)
dt (ms)
v (mV)
}
ASSIGNED {
inmda (nA)
iampa (nA)
gnmda (nS)
gampa (nS)
}
STATE {
A (nS)
B (nS)
C (nS)
D (nS)
}
INITIAL {
gnmda=0
gampa=0
A=0
B=0
C=0
D=0
}
BREAKPOINT {
LOCAL count
SOLVE state METHOD cnexp
gnmda=(A-B)/(1+n*exp(-gama*v) )
gampa=(C-D)
inmda =(1e-3)*gnmda*(v-e)
iampa= (1e-3)*gampa*(v-e)
}
NET_RECEIVE(weight_ampa, weight_nmda) {
INITIAL {
gampamax = weight_ampa
gnmdamax = weight_nmda
}
gampamax = weight_ampa
gnmdamax = weight_nmda
A = A+ gnmdamax
B = B+ gnmdamax
C = C+ gampamax
D = D+ gampamax
VERBATIM
printf("***********\n");
//printf("A = %.2f\n", A);
//printf("B = %.2f\n", B);
//printf("C = %.2f\n", C);
//printf("D = %.2f\n", D);
ENDVERBATIM
}
DERIVATIVE state {
A'=-A/tau1
B'=-B/tau2
C'=-C/tau3
D'=-D/tau4
}