: Based on Schild et al 1994
: Sodium Calcium Exchanger
: Coded and modified by Nathan Titus 2018
: Development Notes: cao, nao, and ko are actually
: concentrations in the perineuronal space
:
:
NEURON {
SUFFIX nacx
USEION na READ nai,nao WRITE ina
USEION ca READ cao,cai WRITE ica
RANGE ina,ica, icana
RANGE knaca, dnaca, R, F, gbar
}
UNITS {
(molar) = (1/liter) : moles do not appear in units
(mM) = (millimolar)
(uA) = (microamp)
(mA) = (milliamp)
(mV) = (millivolt)
(um) = (micron)
}
PARAMETER {
knaca = 36e-9 :(uA/(mM^4))
dnaca = .0036 :(1/(mM^4))
R = 8314 (mV)
F = 96500
gbar = 316 (1/cm2) :converts from uA to mA/cm^2, 3 uA/cm^2 for cai=.0002 and cao=2
}
ASSIGNED {
v (mV)
nai (mM)
nao (mM)
cai (mM)
cao (mM)
ina (mA/cm2)
ica (mA/cm2)
celsius (degC)
icana (mA/cm2)
inaca (mA/cm2)
}
INITIAL {
rates(v)
}
BREAKPOINT {
rates(v)
icana = inaca
ina = 3*inaca
ica = -2*inaca
}
UNITSOFF
FUNCTION rates(Vm (mV)) {
LOCAL q10, T, dfcain, dfcaout, s
T = 273 + celsius
q10 = (2.2*(T-296.0)+(310.0-T))/14.0
dfcain = nai^3*cao*exp(0.5*Vm*F/(R*T))
dfcaout = nao^3*cai*exp(-0.5*Vm*F/(R*T))
s = 1 + dnaca*(cai*nao^3 + cao*nai^3)
inaca = gbar*q10*knaca*(dfcain-dfcaout)/s
}
UNITSON