TITLE Slow Ca-dependent cation current
:
: We've moved to the Model described by Nillus in 2004, while keeping the description of the nanodomain
: modified by Canavier too include separate pool for ICan calcium microdomain
: at this point ican doesn't activate other pools of calcium need to declare a new ion species
: because at this point the code requires pools for SK+BK+T inactivation and ICAN that decay at different rates
INDEPENDENT {
t FROM 0 TO 1 WITH 1 (ms)
}
NEURON {
SUFFIX icannoND
USEION ca READ cai
RANGE erev
RANGE gbar,itrpm4
RANGE beta,taumin
NONSPECIFIC_CURRENT itrpm4
}
UNITS {
(mA)=(milliamp)
(mV)=(millivolt)
(molar)=(1/liter)
(mM)=(millimolar)
(um)=(micron)
(msM)=(ms mM)
FARADAY=(faraday) (coulomb)
}
PARAMETER {
v (mV)
erev=0 (mV) : reversal potential
cai (mM) : will now decay to bulk cai
gbar=0.0001 (mho/cm2)
: middle point of activation fct, for ip3 as somacar, for current injection
taumin=0.1 (ms) : minimal value of time constant
Kd = 87e-3 (mM) :20e-3
}
STATE {
can (mM)
Po
}
ASSIGNED {
jip3p (mM/ms)
ican (mA/cm2)
itrpm4 (mA/cm2)
Po_inf
Tau (ms)
:cai (mM)
}
BREAKPOINT {
SOLVE states METHOD derivimplicit
itrpm4=gbar*Po*(v-erev)
}
DERIVATIVE states {
evaluate_fct(v,cai)
Po'=(Po_inf-Po)/Tau
}
FUNCTION MyExp(x) {
if (x<-50) {MyExp=0}
else if (x>50) {MyExp=exp(50)}
else {MyExp=exp(x)}
}
UNITSOFF
INITIAL {
: activation kinetics are assumed to be at 22 deg. C
: Q10 is assumed to be 3
evaluate_fct(v,cai)
}
PROCEDURE evaluate_fct(v(mV),cai(mM)) {
LOCAL alpha, alpha2, beta
alpha=0.0057*MyExp(0.0060*v)
beta=0.033*MyExp(-0.019*v)
alpha2=alpha/(1+(Kd/cai))
Po_inf=alpha2/(alpha2+beta)
Tau=1/(alpha2+beta)
if (Tau<taumin) {Tau=taumin} : min value of time cst
}
UNITSON