COMMENT
26 Ago 2002 Modification of original channel to allow variable time step
and to correct an initialization error.
Done by Michael Hines (michael.hines@yale.edu) and Ruggero Scorcioni (rscorcio@gmu.edu)
at EU Advance Course in Computational Neuroscience. Obidos, Portugal
km.mod
Potassium channel, Hodgkin-Huxley style kinetics
Based on I-M (muscarinic K channel)
Slow, noninactivating
Original Author: Zach Mainen, Salk Institute, 1995, zach@salk.edu
ENDCOMMENT
INDEPENDENT {t FROM 0 TO 1 WITH 1 (ms)}
NEURON {
SUFFIX km
USEION k READ ek WRITE ik
RANGE n, gk, gbar
RANGE ninf, ntau
GLOBAL Ra, Rb
GLOBAL q10, temp, tadj, vmin, vmax
}
UNITS {
(mA) = (milliamp)
(mV) = (millivolt)
(pS) = (picosiemens)
(um) = (micron)
}
PARAMETER {
: 0.03 mho/cm2
gbar = 10 (pS/um2)
v (mV)
: v 1/2 for inf
tha = -30 (mV)
: inf slope
qa = 9 (mV)
: max act rate (slow)
Ra = 0.001 (/ms)
: max deact rate (slow)
Rb = 0.001 (/ms)
dt (ms)
celsius (degC)
: original temp
temp = 23 (degC)
: temp sensitivity
q10 = 2.3
vmin = -120 (mV)
vmax = 100 (mV)
}
ASSIGNED {
a (/ms)
b (/ms)
ik (mA/cm2)
gk (pS/um2)
ek (mV)
ninf
ntau (ms)
tadj
}
STATE {
n
}
INITIAL {
trates(v)
n = ninf
}
BREAKPOINT {
SOLVE states METHOD cnexp
gk = tadj * gbar * n
ik = (1e-4) * gk * (v - ek)
}
LOCAL nexp
: Computes state variable n at the current v and dt.
DERIVATIVE states {
trates(v)
n' = (ninf - n) / ntau
}
: Computes rate and other constants at current v.
: Call once from HOC to initialize inf at resting v.
PROCEDURE trates(v) {
TABLE ninf, ntau
DEPEND celsius, temp, Ra, Rb, tha, qa
FROM vmin TO vmax WITH 199
: not consistently executed from here if usetable_hh == 1
rates(v)
: tinc = -dt * tadj
: nexp = 1 - exp(tinc/ntau)
}
: Computes rate and other constants at current v.
: Call once from HOC to initialize inf at resting v.
PROCEDURE rates(v) {
a = Ra * (v - tha) / (1 - exp(-(v - tha) / qa))
b = -Rb * (v - tha) / (1 - exp((v - tha) / qa))
tadj = q10^((celsius - temp) / 10)
ntau = 1/tadj/(a+b)
ninf = a/(a+b)
}