: nav1p9.mod is the NaV1.9 Na+ current from
: Herzog, Cummins, and Waxman 2001 p1353
: This current is also called the ttx-rp current
: (the tetrodotoxin resistent persistant current)
: implemented by Tom Morse version 2/25/07
NEURON {
SUFFIX nav1p9
NONSPECIFIC_CURRENT i
RANGE gbar, ena, slow_inact, m, h, s, gate
RANGE tau_m, tau_h, tau_s
: if slow_inact=1 then ultra-slow inactivation is included
}
UNITS {
(S) = (siemens)
(mV) = (millivolts)
(mA) = (milliamp)
}
PARAMETER {
gbar = 0.0069005 (S/cm2)
ena=62.94 (mV)
A_am9 = 1.032 (/ms): 1.548 in Baker '05 : A for alpha m(9 etc ...)
B_am9 = 6.99 (mV) : -11.01 in Baker '05
C_am9 = -14.87115: -14.871 (mV) in Baker '05
A_ah9 = 0.06435 (/ms) : 0.2574 (/ms) in Baker '05, A for alpha h
B_ah9 = 73.26415 (mV) : 63.264 in Baker '05
C_ah9 = 3.71928 (mV) : 3.7193 in Baker '05
A_as9 = 0.00000016 (/ms) : contributes to ultra slowness
B_as9 = 0 (mV)
gate = 0 (mV)
C_as9 = 12 (mV)
A_bs9 = 0.0005 (/ms)
B_bs9 = 32 (mV)
C_bs9 = 23 (mV)
A_bm9 = 5.79 (/ms) : 8.685 in Baker '05 : A for beta m
B_bm9 = 130.4 (mV) : -112.4 in Baker '05
C_bm9 = 22.9 (mV) : 22.9 in Baker '05
A_bh9 = 0.13496 : 0.53984 in Baker '05 : A for beta h
B_bh9 = 10.27853 : 0.27853 in Baker '05
C_bh9 = -9.09334 : -9.0933 in Baker '05
slow_inact = 1 (1) : to turn on ultra slow inactivation
}
ASSIGNED {
v (mV) : NEURON provides this
i (mA/cm2)
g (S/cm2)
tau_h (ms)
tau_m (ms)
tau_s (ms)
minf
hinf
sinf
}
STATE { m h s}
BREAKPOINT {
SOLVE states METHOD cnexp
g = gbar * m * h * s
i = g * (v-ena)
}
INITIAL {
rates(v) : set time constants and infinity values
: assume that equilibrium has been reached
m = minf
h = hinf
s = sinf
}
DERIVATIVE states {
rates(v)
m' = (minf - m)/tau_m
h' = (hinf - h)/tau_h
s' = (sinf - s)/tau_s
}
FUNCTION alpham(Vm (mV)) (/ms) {
alpham=A_am9/(1+exp((Vm+B_am9)/C_am9))
}
FUNCTION alphah(Vm (mV)) (/ms) {
alphah=A_ah9/(1+exp((Vm+B_ah9)/C_ah9))
}
FUNCTION alphas(Vm (mV)) (/ms) {
alphas=A_as9*exp(-(Vm+gate+B_as9)/C_as9)
}
FUNCTION betam(Vm (mV)) (/ms) {
betam=A_bm9/(1+exp((Vm+B_bm9)/C_bm9))
}
FUNCTION betah(Vm (mV)) (/ms) {
betah=A_bh9/(1+exp((Vm+B_bh9)/C_bh9))
}
FUNCTION betas(Vm (mV)) (/ms) {
betas=A_bs9/(1+exp(-(Vm+gate+B_bs9)/C_bs9))
}
FUNCTION rates(Vm (mV)) (/ms) {
tau_m = 1.0 / (alpham(Vm) + betam(Vm))
minf = alpham(Vm) * tau_m
tau_h = 1.0 / (alphah(Vm) + betah(Vm))
hinf = alphah(Vm) * tau_h
if (slow_inact) {
tau_s = 1.0 / (alphas(Vm) + betas(Vm))
sinf = alphas(Vm) * tau_s
} else {
tau_s = 0.1 : in a tenth of a millisecond we move to within
sinf = 1.0 : 1/e factor towards s = 1
}
}