TITLE Low threshold calcium current
: This is a modification of IT2.mod taken from
: the Lytton et al. model in ModelDB
: accession number 9889 but which is from other sources (see below)
: to model the I_CaT in Liu et al. 1998
: Tom M Morse 20070803
: Ca++ current responsible for low threshold spikes (LTS)
: RETICULAR THALAMUS
: Differential equations
:
: Model of Huguenard & McCormick, J Neurophysiol 68: 1373-1383, 1992.
: The kinetics is described by standard equations (NOT GHK)
: using a m2h format, according to the voltage-clamp data
: (whole cell patch clamp) of Huguenard & Prince, J Neurosci.
: 12: 3804-3817, 1992. The model was introduced in Destexhe et al.
: J. Neurophysiology 72: 803-818, 1994.
: See http://www.cnl.salk.edu/~alain , http://cns.fmed.ulaval.ca
:
: ACTIVATION FUNCTIONS FROM EXPERIMENTS (NO CORRECTION)
:
: Reversal potential taken from Nernst Equation
:
: Written by Alain Destexhe, Salk Institute, Sept 18, 1992
:
INDEPENDENT {t FROM 0 TO 1 WITH 1 (ms)}
NEURON {
SUFFIX cat
USEION ca READ cai WRITE ica
POINTER gbar
RANGE m_inf, tau_m, h_inf, tau_h, shift, i, carev
}
UNITS {
(molar) = (1/liter)
(mV) = (millivolt)
(mA) = (milliamp)
(mM) = (millimolar)
(S) = (siemens)
FARADAY = (faraday) (coulomb)
R = (k-mole) (joule/degC)
}
PARAMETER {
v (mV)
: celsius = 36 (degC)
: eca = 120 (mV)
gbar (S/cm2) := .00175 (mho/cm2)
: shift = 2 (mV) : screening charge for Ca_o = 2 mM
cai (mM) : = 2.4e-4 (mM) : adjusted for eca=120 mV
: p.2319 Liu et al. 1998
cao = 3 (mM) : original 2 (mM)
}
STATE {
m h
}
ASSIGNED {
ica (mA/cm2)
i (mA/cm2)
carev (mV)
m_inf
tau_m (ms)
h_inf
tau_h (ms)
celsius (degC)
}
BREAKPOINT {
SOLVE castate METHOD cnexp
carev = (1e3) * (R*(celsius+273.15))/(2*FARADAY) * log (cao/cai)
ica = gbar * m*m*m*h * (v-carev)
i = ica
}
DERIVATIVE castate {
evaluate_fct(v)
m' = (m_inf - m) / tau_m
h' = (h_inf - h) / tau_h
}
UNITSOFF
INITIAL {
evaluate_fct(v)
m = m_inf
h = h_inf
}
PROCEDURE evaluate_fct(v(mV)) {
m_inf= 1.0 / (1+exp( -(v+27.1)/7.2 ))
h_inf= 1.0 / (1+exp( (v+32.1)/5.5 ))
tau_m = 21.7 - 21.3 / (1+exp( -(v+68.1)/20.5 ))
tau_h = 105 - 89.8 / (1+exp( -(v+55)/16.9 ))
}
UNITSON