% Model for action potential of skeletal muscle fiber % Adrian RH, Chandler WK, Hodgkin AL. Voltage clamp experiments in striated muscle % fibres. J Physiol 208:607-44, 1970. % implemented by Dr. Sheng-Nan Wu % Initial values Initial Vm=-95, Vt=-95, m=0.0, h=1.0, n=0.0 % Stimulus Param period=200, iStim_mag=3, iStim_beg=1, iStim_dur=0.5 iStim= iStim_mag * heav(mod(t,period)-iStim_beg) * heav(iStim_beg+iStim_dur-mod(t,period)) % Values of the model parameters Param gNa_max=1.0, gK_max=0.415, gL_max=0.0024 Param ENa=50.0, EK=-70.0, EL=-95.0, Cm=0.0090, Ct=0.04, Rs=15.0 Param alpha_m_max=0.208 Param beta_m_max=2.081 Param alpha_n_max=0.0229 Param beta_n_max=0.09616 Param alpha_h_max=0.0156 Param beta_h_max=3.382 Param Em=-42.0, En=-40.0, Eh=-41.0 Param v_alpha_m=10.0, v_alpha_h=14.7, v_alpha_n=7.0 Param v_beta_m=18.0, v_beta_h=7.6, v_beta_n=40.0 % Expressions Ina= (gNa_max * m * m * m * h * (Vm - ENa)) beta_n= (beta_n_max * exp(((En - Vm) / v_beta_n))) beta_m= (beta_m_max * exp(((Em - Vm) / v_beta_m))) beta_h= (beta_h_max / (1.0 + exp(((Eh - Vm) / v_beta_h)))) IT= ((Vm - Vt) / Rs) IL= (gL_max * (Vm - EL)) IK= (gK_max * n * n * n * n * (Vm - EK)) alpha_n= (alpha_n_max * (Vm - En) / (1.0 - exp(((En - Vm) / v_alpha_n)))) alpha_m= (alpha_m_max * (Vm - Em) / (1.0 - exp(((Em - Vm) / v_alpha_m)))) alpha_h= (alpha_h_max * exp(((Eh - Vm) / v_alpha_h))) % Differential equations dVm/dt= ((iStim - (INa + IK + IL + IT)) / Cm) dm/dt= ((alpha_m * (1.0 - m)) - (beta_m * m)) dh/dt= ((alpha_h * (1.0 - h)) - (beta_h * h)) dn/dt= ((alpha_n * (1.0 - n)) - (beta_n * n)) dVt/dt=((Vm - Vt) / (Rs * Ct)) % Numerical and plotting parameters for xpp @ meth=Euler, dt=0.01, total=25 @ yp=v, yhi=50, ylo=-110, xlo=0, xhi=25, bounds=5000 done