COMMENT
Biexponential single synaptic activation that occurs at a user-specified time.
The conductance varies with membrane potential to emulate NMDA-gated conductance.
Based on syn_g.mod
ENDCOMMENT
NEURON {
POINT_PROCESS nmda
RANGE onset, tau0, tau1, gmax, e, i, g
NONSPECIFIC_CURRENT i
}
UNITS {
(nA) = (nanoamp)
(mV) = (millivolt)
(umho) = (micromho)
}
PARAMETER {
onset=2 (ms)
tau0=3 (ms)
tau1=90 (ms)
gmax=0.001 (umho)
e=5 (mV)
v (mV)
nmg = 0.3 (1) : eta*[Mg]o for eta = 0.25 (/mM) and [Mg]o = 1 mM
: 0.33 used in Polsky's glutamate.mod
: Zador et al. 1990 assumed eta = 0.33 (/mM)
gamma = 0.08 (/mV) : .08 used in Polsky's glutamate.mod
: Zador et al. 1990 used 0.06 (/mV)
}
ASSIGNED { i (nA) g (umho) }
LOCAL a[2]
LOCAL tpeak
LOCAL adjust
LOCAL amp
BREAKPOINT {
if (gmax) {
at_time(onset) : tell integrator when g starts to change
g = cond(t)
}
i = g*(v - e)
}
FUNCTION cond(x(ms))(umho) {
tpeak=tau0*tau1*log(tau0/tau1)/(tau0-tau1)
adjust=1/((1-exp(-tpeak/tau0))-(1-exp(-tpeak/tau1)))
amp=adjust*gmax
if (x < onset) {
cond = 0
}
else {
a[0]=1-exp(-(x-onset)/tau0)
a[1]=1-exp(-(x-onset)/tau1)
: cond = amp*(a[0]-a[1])
cond = (amp*(a[0]-a[1])/(1 + nmg*exp(-gamma*(v))))
}
}