/* =========================================================== Author: Daniel Keller e-mail: daniel.keller@epfl.ch Institution: Ecole Polytechnique Federale de Lausanne Reference: Keller D, Babai N, Kochubey O, Han Y, Markram H, Schürmann F, et al. (2015) An Exclusion Zone for Ca2+ Channels around Docked Vesicles Explains Release Control by Multiple Channels at a CNS Synapse. PLoS Comput Biol 11(5): e1004253. doi:10.1371/journal.pcbi.1004253 ============================================================= */ NOTIFICATIONS { VARYING_PROBABILITY_REPORT=FALSE } //for AP titration, select as needed /*sprintf(data_header_str,"../data_files/linearap0.1/") sprintf(data_header_str,"../data_files/linearap0.2/") sprintf(data_header_str,"../data_files/linearap0.4/") sprintf(data_header_str,"../data_files/linearap0.6/") sprintf(data_header_str,"../data_files/linearap0.8/")*/ sprintf(data_header_str,"../data_files/linearap1.0/") /*sprintf(data_header_str,"../data_files/linearap20/") sprintf(data_header_str,"../data_files/linearap40/") sprintf(data_header_str,"../data_files/linearap80/")*/ sprintf(seed_str,"%04g",SEED) INCLUDE_FILE="../../rates/rates.mdl" INCLUDE_FILE="variables.mdl" TIME_STEP = dt ITERATIONS = it EFFECTOR_GRID_DENSITY = grid_density ACCURATE_3D_REACTIONS = TRUE MICROSCOPIC_REVERSIBILITY=TRUE INCLUDE_FILE="../../parameters/geometry_parameters.mdl" INCLUDE_FILE="../../parameters/epspstart.mdl" SPACE_STEP= 0.001 ITERATIONS = it EFFECTOR_GRID_DENSITY = grid_density INTERACTION_RADIUS=0.002 PARTITION_X=[[-dendrite_length/2-0.05 TO dendrite_length/2+0.05 STEP 0.03]] PARTITION_Y=[[-dendrite_width/2-0.05 TO dendrite_width/2+0.05 STEP 0.03]] PARTITION_Z=[[-2.0001 TO 2.0001 STEP 0.03]] /* --------- Mechanisms ------------- */ INCLUDE_FILE="../../molecules/molecules.mdl" INCLUDE_FILE="../../mechanisms/mechanisms.mdl" /* ----- Define Physical Objects ------- */ INCLUDE_FILE="../../geometry_files/bbp_sphere2.mdl" INCLUDE_FILE="../../geometry_files/sampling_boxes.mdl" INCLUDE_FILE="../../geometry_files/cbp_sheets.mdl" INCLUDE_FILE="../../release_events_files/glutamate_release.mdl" INCLUDE_FILE="../../geometry_files/bouton2.mdl" INCLUDE_FILE="../../geometry_files/bouton_sensor.mdl" INSTANTIATE tissue OBJECT { sphere1 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ -0.0992 , 0.0267 , 0.0435+0.5*gap_size] } sphere2 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ -0.0992 , 0.0965 , 0.0834+0.5*gap_size] } sphere3 OBJECT sphere1 { SCALE=[2.0,2.0,2.0] TRANSLATE=[ -0.0496 , 0.0447 , 0.0223+0.5*gap_size] } sphere4 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ -0.0496 , -0.0079 , 0.0304+0.5*gap_size] } sphere5 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ -0.0496 , 0.0154 , 0.0700+0.5*gap_size] } sphere6 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ -0.0496 , 0.0811 , 0.0723+0.5*gap_size] } sphere7 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , 0.0528 , 0.0328+0.5*gap_size] } sphere8 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , -0.0003 , 0.0324+0.5*gap_size] } sphere9 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , 0.1101 , 0.0669+0.5*gap_size] } sphere10 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , 0.0600 , 0.0706+0.5*gap_size] } sphere11 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , -0.0818 , 0.0802+0.5*gap_size] } sphere12 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , -0.0475 , 0.0639+0.5*gap_size] } sphere13 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0 , 0.1101 , 0.0797+0.5*gap_size] } sphere14 OBJECT sphere1 { SCALE=[2.0,2.0,2.0] TRANSLATE=[ 0.0496 , 0.0151 , 0.0227+0.5*gap_size] } sphere15 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0496 , -0.0299 , 0.0289+0.5*gap_size] } sphere16 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0496 , 0.0140 , 0.0791+0.5*gap_size] } sphere17 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0496 , -0.0329 , 0.0714+0.5*gap_size] } sphere18 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0992 , 0.0140 , 0.0864+0.5*gap_size] } sphere19 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0992 , 0.0537 , 0.0361+0.5*gap_size] } sphere20 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0992 , -0.0908 , 0.0529+0.5*gap_size] } sphere21 OBJECT sphere1 { SCALE=[2.5,2.5,2.5] TRANSLATE=[ 0.0992 , 0.1107 , 0.0793+0.5*gap_size] } bouton OBJECT bouton {} bouton_box OBJECT bouton_box {} bouton_box_top OBJECT bouton_box_top {} boutonactivezone OBJECT boutonactivezone {} Glu_release_site_1 OBJECT Glu_release_site_1 {} RELEASE_active RELEASE_SITE { SHAPE=LIST MOLECULE_POSITIONS { Ca_HH.C0' [ -0.0368 , -0.0711,0.5*gap_size] Ca_HH.C0' [ -0.1133 , 0.0171,0.5*gap_size] Ca_HH.C0' [ 0.0866 , -0.0728,0.5*gap_size] Ca_HH.C0' [ 0.1075 , -0.0627,0.5*gap_size] Ca_HH.C0' [ 0.0941 , 0.0768,0.5*gap_size] Ca_HH.C0' [ -0.0062 , -0.0963,0.5*gap_size] Ca_HH.C0' [ 0.0998 , 0.0235,0.5*gap_size] Ca_HH.C0' [ 0.1189 , 0.0999,0.5*gap_size] Ca_HH.C0' [ 0.0869 , -0.0742,0.5*gap_size] Ca_HH.C0' [ 0.1100 , 0.0280,0.5*gap_size] Ca_HH.C0' [ 0.0886 , 0.1074,0.5*gap_size] Ca_HH.C0' [ -0.0248 , 0.0972,0.5*gap_size] Ca_HH.C0' [ 0.0972 , -0.0503,0.5*gap_size] Ca_HH.C0' [ -0.0639 , 0.1029,0.5*gap_size] } SITE_DIAMETER = 0.005 } RELEASE_boutonsensoraz0 CUBIC_RELEASE_SITE { LOCATION = [ -0.0496 , 0.0447 ,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor0.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_boutonsensoraz1 CUBIC_RELEASE_SITE { LOCATION = [ -0.0496 , -0.0079,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor1.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_boutonsensoraz2 CUBIC_RELEASE_SITE { LOCATION = [ 0 , 0.0528 ,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor2.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_boutonsensoraz3 CUBIC_RELEASE_SITE { LOCATION = [ 0 , -0.0003 ,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor3.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_boutonsensoraz4 CUBIC_RELEASE_SITE { LOCATION = [ 0.0496 , 0.0151 ,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor4.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_boutonsensoraz5 CUBIC_RELEASE_SITE { LOCATION = [ 0.0496 , -0.0299,0.5*gap_size+1.5 *fuzz+0.002] LIGAND = release_sensor5.a.x0 NUMBER_TO_RELEASE = 1 SITE_DIAMETER = [0.001,0.001,0.001] } RELEASE_pumps RELEASE_SITE { SHAPE = tissue.bouton.bouton0[topandbottom] MOLECULE = generic_pump.U' NUMBER_TO_RELEASE=0 } RELEASE_ATP.CaB RELEASE_SITE { SHAPE = tissue.bouton_box_top[ALL] MOLECULE = ATP.CaB CONCENTRATION=atp_concentration * bound_atp_fraction } RELEASE_ATP.MgB RELEASE_SITE { SHAPE = tissue.bouton_box_top[ALL] MOLECULE = ATP.MgB CONCENTRATION=atp_concentration * mgbound_atp_fraction } RELEASE_ATP.U RELEASE_SITE { SHAPE = tissue.bouton_box_top[ALL] MOLECULE = ATP.U CONCENTRATION=atp_concentration * unbound_atp_fraction } RELEASE_EGTA.B RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = EGTA.B CONCENTRATION=EGTA_concentration * EGTA_bound_fraction } RELEASE_EGTA.U RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = EGTA.U CONCENTRATION=EGTA_concentration * EGTA_unbound_fraction } RELEASE_BAPTA.B RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = BAPTA.B CONCENTRATION=BAPTA_concentration * BAPTA_bound_fraction } RELEASE_BAPTA.U RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = BAPTA.U CONCENTRATION=BAPTA_concentration * BAPTA_unbound_fraction } RELEASE_bouton_slowB RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = slow_sp.B CONCENTRATION=slow_cbp_concentration * bound_cbp_fraction } RELEASE_bouton_slowU RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = slow_sp.U CONCENTRATION=slow_cbp_concentration * unbound_cbp_fraction } RELEASE_bouton_parvalbuminB RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = parvalbumin.B CONCENTRATION=parvalbumin_concentration * bound_parvalbumin_fraction } RELEASE_bouton_parvalbuminU RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = parvalbumin.U CONCENTRATION=parvalbumin_concentration * unbound_parvalbumin_fraction } RELEASE_bouton_parvalbuminM RELEASE_SITE { SHAPE = tissue.bouton.bouton0[ALL] MOLECULE = parvalbumin.M CONCENTRATION=parvalbumin_concentration * mgbound_parvalbumin_fraction } } REACTION_DATA_OUTPUT { OUTPUT_BUFFER_SIZE=10 STEP = freq { COUNT[ Ca_HH.O,WORLD] } => "Cachan.O." & seed_str { COUNT[ release_sensor0.a.x6, WORLD]+ COUNT[ release_sensor0.b.x6, WORLD]+ COUNT[ release_sensor0.c.x6, WORLD]+ COUNT[ release_sensor0.d, WORLD] } => "x0." & seed_str { COUNT[ release_sensor1.a.x6, WORLD]+ COUNT[ release_sensor1.b.x6, WORLD]+ COUNT[ release_sensor1.c.x6, WORLD]+ COUNT[ release_sensor1.d, WORLD] } => "x1." & seed_str { COUNT[ release_sensor2.a.x6, WORLD]+ COUNT[ release_sensor2.b.x6, WORLD]+ COUNT[ release_sensor2.c.x6, WORLD]+ COUNT[ release_sensor2.d, WORLD] } => "x2." & seed_str { COUNT[ release_sensor3.a.x6, WORLD]+ COUNT[ release_sensor3.b.x6, WORLD]+ COUNT[ release_sensor3.c.x6, WORLD]+ COUNT[ release_sensor3.d, WORLD] } => "x3." & seed_str { COUNT[ release_sensor4.a.x6, WORLD]+ COUNT[ release_sensor4.b.x6, WORLD]+ COUNT[ release_sensor4.c.x6, WORLD]+ COUNT[ release_sensor4.d, WORLD] } => "x4." & seed_str { COUNT[ release_sensor5.a.x6, WORLD]+ COUNT[ release_sensor5.b.x6, WORLD]+ COUNT[ release_sensor5.c.x6, WORLD]+ COUNT[ release_sensor5.d, WORLD] } => "x5." & seed_str { COUNT[ io_hh,WORLD] } => "ica." & seed_str }